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BEHAVIOUR publishes original research pursuing Tinbergenâs four questions and questions resulting from the interrelationship among the four. In addition, the editorial board encourages reviews of behavioural biology that illuminate emergent trends and new directions in behavioural research. Niko Tinbergen (1907-1988) defined 4 questions for research in behavioral biology: Proximate causation of behaviour: 1. How does an animal use its sensory and motor abilities to activate and modify its behaviour patterns? (physiological mechanisms) 2. How does an animalâs behaviour change during its growth, especially in response to the experiences that it has while maturing? (ontogeny of development) Ultimate causation of behaviour
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